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Visit the Office of Graduate Education for info on our many doctoral programs and the graduate education application process at the School of Medicine.

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In 1956, the School of Medicine became the first school to offer combined training for an M.D. and a Ph.D. in the basic medical sciences. Among the graduates of this program are 1994 Nobel Prize recipient Alfred G. Gilman, M.D., Ph.D., 1998 Nobel Prize recipient Ferid Murad, M.D., Ph.D., and former U.S. Surgeon General David Satcher, M.D., Ph.D. Today, the school is one of only 43 institutions nationwide to offer the dual degrees through the National Institutes of Health (NIH)-funded Medical Scientist Training Program (MSTP).

MSTP students at the School of Medicine earn their medical degrees through the University Program of the School of Medicine and their Ph.D.s in the basic science field of their choice through the School of Graduate Studies. Currently, Ph.D. programs are offered in Biochemistry; Biomedical Engineering; Cell Biology; Clinical Translational Science; Epidemiology & Biostatistics; Genetics & Genome Sciences; Molecular Biology & Microbiology; Molecular Virology; Neurosciences; Nutrition; Pathology (with tracks in Cancer Biology, Immunology, or Molecular & Cellular Basis of Disease); Pharmacology; Physiology & Biophysics; and Systems Biology & Bioinformatics. Additional training programs within these Ph.D. areas span other specific fields, such as Developmental Biology.

Students begin the MSTP by attending medical school for two years. During this time, they are able to complete most of their Ph.D. coursework as well as the laboratory rotations necessary for them to choose their Ph.D. concentrations and mentors. Students then spend an average of  four years conducting research and preparing a thesis to satisfy the requirements for the Ph.D. They then finish the last two years of the medical school program.